Early Civilizations:
Complex Societies of the New
and Old Worlds
Dr. Kathryn Denning
Anth 2150, Sept 2007 - Apr 2008
6 Nov 2007...
Welcome back!
Plan for the day
1 Announcements.
QUIZ #2 will be held on November 7th in tutorial. Review guidelines are here. The quiz will cover material from Oct 16, 23, 30... it will be primarily (but not exclusively) drawn from the textbook, chapters 4, 5, 6, 7.
After that, you'll be watching a short film clip about our hominid ancestors.
Next week in tutorial -- you'll see, close-up, various hominid skulls
2 Lecture
- a little recap/review of archaeological methods
- the story of humanity's journey begins: Meet the Ancestors!
- including film clips from The Story of Lucy (York vid #1033) and the BBC feature Walking with Cavemen
For the first part of this course....
We've focused on how archaeology is done
Ch 1 - the early days of archaeology, the major developments of prehistory, the past in the present
Ch 2 - the record of the past - what the archaeological record is / different kinds of site preservation.... how we reconstruct ancient lifeways, explain cultural change, and do archaeological research (plan, acquire data, analyse, interpret, publish)... the importance of context... ethical responsibilities...
Ch 3 - acquiring the record - how do we actually find sites and excavate? How do we determine dates?
Ch 4 - how did people live? - ancient technologies (stone, bone, metal, wood, ceramics, basketry/textiles) and how we analyse them... how did people subsist? and analysis of ancient foods... animals and plants... reconstructing ancient diet
Ch 5 - individuals and interactions - understanding things like social ranking, gender, ethnicity, inequality, trade/exchange... e.g. Otzi, division of labour by gender at Abu Hureyra, plantations with enslaved people, war casualties in ancient Egypt, ancient exchange networks e.g. obsidian sourcing, Uluburun ship
Ch 6 - studying the intangible - the framework of common beliefs of ancient civilizations, ways of interpreting ancient art, the archaeology of death, the importance of context in interpreting artifacts, sacred places and their symbolism (e.g. ancient astronomical alignments)
Ch 7 - explaining the past - culture history, cultural change (e.g. cultural selection, diffusion, migration), doing archaeology by observation / experiment (ethnoarchaeology, experimental archaeology).... Explaining cultural change (different theoretical models, like cultural ecology, multilinear cultural evolution, processual and post-processual archaeology) ...
- and we had a week on bioarchaeological methods, or how researchers obtain information from the skeleton
Then Last week...
We took a Hallowe'en detour, to think about the role of the dead in societies, past and present... from the 'undead' vampire to mummies...
And now... we'll begin telling the story of humanity's journey from our earliest human ancestors to civilizations. This is the information we have obtained through the methods of archaeology.
Remember, the four major developments in human prehistory (from Ch 1):
- origins of humankind around 2.5 mya
- origins of archaic humans like Homo erectus, and anatomically modern people, from 1.9 mya to 15 000 bp, and the spread of fully modern humans throughout the world by 15 000 bp
- origins of food production around 11 000 bp
- origins of civilizations around 3100 BC
This week's readings
Chapter 8 - Human Origins
Chapter 9 - African exodus
Ch 8 Topics:
- locating ourselves in time
- primates
- fossil evidence
- australopithecines (e.g. Australopithecus afarensis)
- early Homo (H. habilis)
- early human technology
- foraging, scavenging, hunting
- mind... and development of language
- early social organization
Ch 9 Topics:
- Ice age
- Homo ergaster (Africa)
- Homo erectus (Asia 1.9mya - 200 000 bp)
- archaic H. sapiens (400 000 - 130 000 bp)
- Neanderthals (150 000 - 30 000 bp)
- origins of anatomically modern humans ( 180 000 - 150 000)
- out of Africa
Evolution
Social context of theories about evolution
- We all need explanations for how the world came to be this way - who are we? where do we come from? what does it mean to be human? what is our relationship to the natural world? Every culture has a belief system that addresses these questions.
However, science addresses them in a different way -- through developing theories, formulating hypotheses, gathering data, testing the hypotheses, modifying the theories... in a never-ending cycle which has resulted in a vast increase in our knowledge about evolution.
Highly public debates about what should be taught in schools are frequently more about politics and religion than about understanding biology.
Important principles:
1) just because we don't fully understand a process yet doesn't mean that the only explanation is that a divine being made it happen. Another option is to say that we don't know everything yet, and to keep investigating.
2) the study of evolution does not preclude belief in a divine being. That isn't the problem in the evolution-creation controversy. (The problem is the question of what exactly the divine being - if there is one - has been doing, and how exactly we can know that.)
Locating ourselves in time
What are the key dates for biological evolution? (All dates are approximate!)
1 |
2 |
3 |
4 |
|
Date on a one-year calendar[1] |
Actual Date[2] |
What was new? |
Notes on Extinctions etc. in terms of Geological periods |
GEOLOGICAL PERIODS Only start date is given. Dates are approximate. |
Jan 1 |
15 bya |
the universe formed |
|
PRECAMBRIAN |
Sept 24 |
4.5 bya |
Earth formed |
|
|
Sept 25 |
4 bya |
bacteria appeared |
|
|
Nov 15 |
1.7 bya |
multicellular organisms appeared |
|
|
Dec 17 |
570 mya
|
complex organisms, jellyfish, worms, invertebrates |
Explosion of biodiversity in Cambrian period |
CAMBRIAN 570 MYA |
Dec 19 |
510 mya |
fish, coral, vertebrates |
500 mya (late Cambrian) massive extinction. 440 mya (Ordovician), large extinction. |
ORDOVICIAN 500 MYA
|
Dec 21 |
425 mya |
land plants, insects, jawed fish, air-breathing animals first forests 395 mya |
Around 365 mya, Devonian extinction (70% of species lost) |
SILURIAN 430 MYA
DEVONIAN 395 MYA |
Dec 23 |
350 mya |
amphibians and reptiles (dinosaurs appeared 230 mya) mammal-like reptiles 280 mya egg-laying mammals 225 mya Great Age of Dinosaurs 190 mya toothed birds 190 mya |
Around 245 mya (end Permian extinction), 96% of all species eliminated, including 75% of all vertebrates Around 200 mya (late Triassic extinction) 25% of species eliminated |
CARBONIFEROUS 345 MYA PERMIAN 280 MYA TRIASSIC 225 MYA JURASSIC 190 MYA
|
Dec 25 |
150 mya |
lots of reptiles, fish, amphibians modern birds placental & marsupial mammals |
n.b. continents all one (Pangaea) until splits start ca. 200 mya |
CRETACEOUS 136 MYA |
Dec 26 |
65 mya
|
mammals (warm-blooded or homeothermic, i.e. able to regulate own body temperature, and live in more climates; increased investment in offspring; increased brain development) |
65 mya Cretaceous-Tertiary boundary (mass extinction, 85% of all species eliminated, including dinosaurs) |
TERTIARY 65 MYA Tertiary Epochs, below Paleocene 65mya
|
|
60 mya |
primates (more like prosimians) with grasping hands, opposable thumbs, stereoscopic vision, nails, collarbone, 1-2 offspring at a time |
Weather everywhere is tropical 40 mya: continents near present positions |
Eocene 55 mya Oligocene 34 mya |
Dec 29 |
35 mya |
monkeys and apes (anthropoids): flatter faces, nonmobile ears, very dextrous hands |
33 mya late Eocene extinction, loss of many mammals – global cooling |
Miocene 23 mya |
|
15 mya |
ape-like ancestors of humans (hominoids), e.g. Proconsul |
Miocene |
|
Dec 31, 10:30 pm |
4 mya |
human-like beings (hominids: Australopithecus afarensis and onwards to Homo): bipedal, bigger brain, reduction of face, teeth, jaws |
Pliocene |
Pliocene 5 mya Pleistocene 1.8 mya |
Dec 31 Stone tool use: 11:00:00 pm Agriculture: 11:59:20 Writing: 11:59:51 0 AD: 11:59:56 Now Nuclear weaponry, global culture, space travel |
100 000 ya
anatomically modern humans (Homo sapiens sapiens): larger brain, chin, smaller brow ridges, lighter skeleton, smaller teeth and jaws |
11 000 ya, substantial extinction of large mammals (climate change)
now – the sixth great extinction? |
Holocene 0.01 mya |
[1] This metaphor comes from Carl Sagan, Dragons of Eden, 1977 p15
[2] 1 billion = 1 000 000 000 1 million = 1 000 000 bya = billion years ago mya = million years ago
Locating ourselves within the sweep of biological diversity
binomial nomenclature:
Genus | species | subspecies |
Homo | sapiens | sapiens |
Always write as Homo sapiens. Generally, italicize.
Primate Taxonomy
Domain - Eukarya
Kingdom - Animalia
Phylum - Chordata
Subphylum - Vertebrata
Class - Mammalia (more specifically, eutherians - cf monotremes and marsupials)
Order - Primates
etc.
Primates - our family
Primate Learning and Behavior
Have a greater dependence on flexible, learned behavior.
Tend to live in social groups.
Males are permanent members of many primate social groups, a situation unusual among mammals.
Arboreal Hypothesis
Arboreal (tree) living was the most important factor in the evolution of primates.
Prehensile hand is adapted to climbing in the trees.
A variety of foods led to the omnivorous diet and generalized dentition.
Visual Predation Hypothesis
Primates may have first adapted to shrubby forest undergrowth and the lowest tiers of the forest canopy.
Forward facing eyes enabled primates to judge distance when grabbing for insects.
Flowering plants may have influenced primate evolution.
Primate Habitats
Most live in tropical or semitropical areas of the new and old worlds.
Most are arboreal, living in forest or woodland habitats.
No nonhuman primate is adapted to a fully terrestrial environment; all spend some time in the trees.
Primate Diet and Teeth
Generally omnivorous, reflected in their generalized dentition.
Most eat a combination of fruits, leaves, and insects.
Some primates kill and eat small mammals.
Some primates are dietary specialists for leaves.
Most have four types of teeth: incisors, canines, premolars and molars.
Primate Locomotion
Most primates are quadrupedal, using all four limbs in their locomotion.
Arm swinging is found among the apes.
Siamangs of southeast Asia use this exclusively.
Monkeys that use a combination of leaping and arm swinging are termed semibrachiators.
Prehensile tails, found only among the new world monkeys, are used as an aid to locomotion.
Below: 4 main locomotor patterns, although n.b. primates are flexible and able to move in more than one way (unlike some animals, e.g. dogs, horses are more restricted in their locomotion)
Skeletons are from:
*Terrestrial quadruped (savanna baboon)
*Arboreal (bearded saki) New World - prehensile tail
*Vertical clinger/leaper (indri)
*Brachiator (gibbon)
More detail on the locomotor pattern and its connection to habitat and behaviour:
Terrestrial quadrupeds
Front and hind limbs almost even length (forelimbs 90% of hind limb length), fast runners. Food/environment: tend to live in savanna or open areas, sit on the ground to eat seeds etc. Often tend to sleep in trees or on cliffs to avoid predators. Usually live in large groups. Tail not prehensile.
Savanna baboon
Papio cynocephalus
Arboreal - tend to walk on top of branches more than swinging below them (think squirrels). Forelimbs 70-80% length of hind limbs. (Need more power in the back for jumping than in the front.) If in the New World, often have prehensile tails. In the Old World, often have long tails which are used for counterbalancing, but are not prehensile. Hands and feet are good for grasping.
White-nosed Bearded Saki (Chiroptes albinasus)
Prehensile Tail: interestingly, also seen in other orders of arboreal animals, like lizards, porcupines, opossums...
Spider monkey, right -->
http://primates.com/monkeys/spidermonkey.html
Vertical clingers/leapers
Most often seen in prosimians (lemurs, tarsiers) - clinging to tree trunks. Hands very good for grasping. Live in forest: some eat bugs, tree sap, invertebrates, while others species eat leaves, fruit etc. Tend to live in small colonies, pairs, or to be mostly solitary.
INDRI (Indri indri)
Tarsier: another vertical clinger/leaper. Note pads on hands and huge (immobile!) eyes.
http://primates.com/philippine-tarsier.html (n.b. not actually smallest 'monkey' ... that's the pygmy marmoset, over in the New World)
Brachiators/ arm swingers: apes, including lesser apes (gibbons and siamangs), great apes (chimps, bonobos, gorillas, orangs). Generally longer arms than legs. Can support entire body hanging from one arm. Live in forest: tend to hang from branches and eat fruit.
Gibbons: live in pairs or very small family groups.
(Humans too... our legs are long by comparison, but we still have the suspensory shoulder adaptation.)
Gibbon (Hylobates sp)
N.b. other locomotor variants
Orangs - quadrumanous - i.e. feet and hands all have opposable thumbs! primarily arboreal, use feet just like hands, unbelievably flexible. Eat fruit, tend to live alone or in very small groups.
Gorillas and chimps - knuckle-walkers but also climb trees. Gorillas live in small extended family groups and primarily eat leaves/stems, chimps in somewhat larger groups and primarily eat fruit.
Bipedalism - mostly just us and our ancestors -- the point at which habitual bipedalism emerged is much-discussed -- but bipedalism does occur occasionally in many primates. e.g. the sifaka bounces along bipedally, injured chimps may walk bipedally, and occasionally a monkey will develop the habit too.
Sifaka travelling bipedally http://primates.com/lemurs/sifbab.html
Primate Behavior
Primates exist in a tremendous range of shapes and sizes, from the pygmy marmoset to the gorilla. They also have a host of different behaviour patterns:
-some live in large groups, others in small groups, pairs, or mostly alone
-eat anything from bugs to fruit to other animals
- in some species, males and females look the same, and in others they differ
- live anywhere from snow to tropics
e.g. Japanese Macaque
http://primates.com/snowmonkeys/snowmonkey.html
e.g. Pygmy Marmoset: http://www.rarespecies.org/pygmy2.htm
Photo gallery showing diversity of primates
http://primates.com/
Importance of Primate Behaviour Studies
Modern African apes and humans last shared a common ancestor between 5 and 8 m.y.a. We are closely related, and although the apes are not themselves frozen in time, they can shed light on early hominid behaviour, as well as modern human behaviour
n.b. most primate species are now threatened, many seriously so (Red List:
http://www.iucnredlist.org/, http://www.iucnredlist.org/info/tables/table4a.htmln.b. this list shows trends... and our lack of knowledge
Nonhuman primate behaviour has been systematically studied since the early 20th century (captive baboons, London Zoo), but that was intermittent, and the first wild population studies came later, 1960s. (e.g. ape studies: Fossey, Goodall, Galdikas)
Many species, such as arboreal monkeys, have scarcely been studied at all.
n.b. many have been studied primarily in captivity, and this shapes their behaviour and our understanding of them (e.g. dominance hierarchies in baboons)
n.b. there are serious logistical challenges to some studies in the wild -- e.g. nocturnal, arboreal, very rare, etc. -- meaning that we don`t actually understand many primates very well yet!
n.b. there has also been a tendency to study primates for specific human agendas, rather than for themselves
Like many higher-order vertebrates, primates have both innate and learned behaviour. Behavioural plasticity (capacity to change) is a key primate adaptation.
Reproductive Strategies
Primates produce only a few young in whom they invest a tremendous amount of parental care. (k –selected rather than r-selected)
The young are very dependent upon their mothers and the group.
http://primates.com/orangutans/baby.htm
And young primates (like many young mammals) play a lot
http://primates.com/snowmonkeys/snowball.html
Mothers and Infants
http://www.hedweb.com/gorilfam.htm
The basic social unit among all primates is the female and her infants.
Monkeys raised without a mother were not able to form lasting affectional ties - Harlow.
The mother-infant relationship is often maintained throughout life.
The role of the male varies according to species: some (e.g. callitrichids) are very active parents, whereas others play no particular role.
Others in the social group may also engage in parenting behaviour, e.g. older siblings, other relatives.
Nonhuman Primate
Cultural Behavior
Cultural behavior is learned and passed from one generation to the next.
Chimpanzee culture includes tools such as termite fishing sticks and leaf sponges.
http://primates.com/chimps/chimpwat.htm
Chimpanzee tool use:
http://news.nationalgeographic.com/news/2004/10/1006_041006_chimps.htmlGorilla tool use:
http://www.livescience.com/animalworld/050929_gorilla_tools.html
Primate Cognitive Abilities
Social interactions and problem-solving abilities demonstrate primate intelligence.
The fact that apes can’t speak has more to do with their anatomy and the language related structures of the brain than intelligence.
They do have communication systems, and our understanding of these is constantly developing.
Primate Sounds:
http://www.monkeymania.co.uk/gosounds/Koko the gorilla:
http://www.gorilla.org/world/daily.Koko meets Mr. Rogers:
http://ali.apple.com/ali_sites/ali/exhibits/1000790/koko_meets_the_stars.htmlhttp://www.pbs.org/teachersource/scienceline/archives/aug99/aug99.shtm
The Primate Continuum
Human brains are larger than primate brains, but the neurological processes are functionally the same.
Nonhuman primates think and have feelings. We may not understand them, but that does not disprove their presence.
An astonishing blindness: That humans are part of an evolutionary continuum is the basis for animal research, yet we cage nonhuman primates with little regard for their needs.
Captive primates should be maintained in social groups and have access to habitat enrichment programs.
photo by Tommy Hayne. Captive monkey in Vietnam, 1994
To proceed with studying hominid evolution, we need to know our way around the skeleton...
Orienting ourselves anatomically
Some key features on a human skeleton: frontal, parietal, mandible, maxilla, occipital, clavicle, ribs, humerus, femur, foramen magnum, incisors, canine, premolars, molars
A few key principles about hominoid & hominid evolution
challenges include these:
When we’re studying fossils: We have to contend with infrequent fossilization, fragmentary remains and the challenges of taphonomy (the changes that occur to organisms or objects after being buried or deposited), including scattering of remains, exposure to the elements, etc. Have to consider why things are found together (deliberate deposition, accidental coincidence, etc.)
- fragmentary, damaged, infrequent fossils, often without a population for comparison ... so it can be hard sometimes to decide whether a fossil is a new species, or a variant on an older one, or whether it's a male or female. Entire skeletons are quite rare.
- this means there's a lot of room for differences of opinion, and different names for the same fossil... usually when a new type of fossil is found, it takes a few years for consensus about it to emerge
- new finds rewrite the taxonomies, quite often... also, you can 'lump' or 'split'
- we can sometimes get DNA out of semi-fossilized bone, but this is rare and only recently possible. Can't get DNA from something completely fossilized. No DNA yet for australopithecines... but they're working on Neanderthal DNA now.
n.b. sometimes there are disputes about DNA analyses.... e.g. what the oldest DNA retrieved is. (Sometimes scientists make announcements but other scientists can't confirm their results)
- dating can be a challenge. Potassium argon dating is the primary method used for dating hominid fossils, but note -- this is dating the volcanic rock layers above or below the fossil, rather than the fossil itself.
[Radioactive potassium decays to argon gas... so we can only use this method on minerals which contain potassium and then trap the argon gas so that we can measure it. Volcanic tuff (compressed ash) has this chemical composition... and there's lots of it in East Africa, which is handy for hominid dating.]
Date range for K-AR - from 250 000 years ago back to the origins of life.
We can also use C14 dating, but that's for later specimens (not fully fossilized), from about 40 000 BP until present.
Note regarding dates/terminology:
mya = million years ago (or, myr ago)
kya = thousand years ago
BP = before present
BC = Before Christ = BCE = Before Common Era
AD = Anno Domini = CE = Common Era
Have a hard time converting these? Think about a timeline...
Also recalling -- how do we picture evolution taking place?
The "March of Progress" iconography... this is not how things happened.
Below, is a better way of representing human evolution.
.. we have many different specimens (these are just a few), and although we can group them approximately with a high degree of confidence, we can't trace a single line of human evolution.
And also recalling... something important about adaptations.
there is variability in any population, and various evolutionary forces then affect the prevalence of different traits. (forces including natural selection, sexual selection, genetic drift, mass extinction events, etc.)
but note! according to standard evolutionary explanations: the variability doesn't arise "for a reason", i.e. there's no big plan or direction for evolution. So, for example, our primate ancestors began to develop nimble hands... but they didn't develop nimble hands so that we could later hold a pencil. They started to develop nimble hands, which proved to be an advantage in a certain environment, and then also turned out to have additional uses later on.
So we didn't evolve nimble hands for the purpose of playing the piano or firing a gun or typing on a keyboard, but it turns out that these are things we can do with our hands.
Similarly, we didn't develop a suspensory shoulder joint so that we could play baseball... it's a left-over feature from our earlier primate heritage. But even though we don't swing through the trees (much) anymore, it came in handy for other things -- like throwing things... rocks, spears, hand grenades, etc.
Human evolution overall
Becoming Human (Flash) http://www.becominghuman.org/
(Anatomy... hominid profiles)
Historical development of Understanding of Human Evolution
It has long been realized that the key trends in human evolution, i.e. the things that separate us from our ape cousins and ancestors, were...
terrestriality - coming down out of the trees
bipedalism - having to walk on two feet (obligate, not occasional)
encephalization - increase in brain size in relation to body, development of language
civilization - complex technology, moral systems, complex society
But in the earliest days of studying human evolution (up to 1920s), it wasn't known which came first...
- there weren't enough fossils, they weren't well understood, and they weren't securely dated
- Arthur Keith 1895 "Only four individuals to represent the millions and millions of men that must have lived and died in Quaternary times!" (German Neanderthal skull, Pithecanthropus from Java, two Cro-Magnon Belgian skulls) (As opposed to now, when we have MANY)
- so different theories were abundant
This is why the Piltdown forgery worked.... people at the time wanted to believe that big brains came before other human traits, so a big-brained 'fossil' with an ape's jaw fit their preconceptions.
Now, we know that's not what happened... Our brains were still pretty small when we became bipedal.